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Papers in Preparation/Review, (*) = Papers with students as first authors

​*Rojas-Pena ML and NV Schizas The Influence of Contrasting Reproductive Modes in the Genetic Population Structure of Caribbean Brittle Stars 

*Jassoud A, MQ Lucas, T Lado-Insua, A Miliou, NV Schizas. The Atlantic Ocean: an impassable barrier for the common octopus Octopus vulgaris.

*Ricaurte M, Schizas NV, Weil E, P Ciborowski, NM Boukli.  Seasonal Temperatures Reveal Unique Markers in the Endangered Coral species Acropora palmata.

Corgosinho PHC, ...NV Schizas. A New Genus and Species of Parastenocarididae Chappuis, 1940 (Copepoda, Harpacticoida) from the Espinhaço Mountain Range (Brazil) with an Unclear Phylogenetic Position within the Fontinalicaridinae Schminke, 2010. Zootaxa

José J. Orozco Juarbe, Paulo Henrique Costa Corgosinho, M Alfaro, NV. Schizas.  Checklist of the Marine Copepods of Puerto Rico.  Caribbean Journal of Science

*Domenech Ramos JP, PHC Corgosinho, NV Schizas Autecology and genetic diversity of Cletocamptus copepods in coastal hypersaline lagoons of Puerto Rico

*Daniel A. Toledo-Rodriguez,  Alex J. Veglia, Nilda M. Jimenez-Marrero, Joyce M. Gomez-Samot,Catherine S. McFadden, Ernesto F. Weil, Nikolaos V. Schizas.  Shadows over Caribbean reefs: Identification of a new invasive soft coral species, Xenia umbellata, in southwest Puerto Rico. bioRxiv preprint doi: (pdf)  

Published Papers or in Press/Accepted. (*) = Papers with students as first authors


*Ricaurte M, Schizas NV, Weil EF, Ciborowski P, Boukli NM. Seasonal Proteome Variations in Orbicella faveolata Reveal Molecular Thermal Stress Adaptations. Proteomes. 2024; 12(3):20.

*Rodriguez-Ferrer G, Appeldoorn RS, Mignucci-Giannoni AA, Rinaldi R, NV Schizas (2024), The presence of two distinct mitochondrial lineages in the bottlenose dolphin (Tursiops truncatus) in Puerto Rico and their affinities with previously reported lineages. Mammalian Biology


Schizas NV, Petersen B, B Rosado (2024) IN MEMORIAM: Govind S. Nadathur (1955-2022). Caribbean Journal of Science 54: 103–106,


*Lebrón-López NE, Nadathur, GS, NV Schizas (2024) Culturable actinobacteria from two marine sponges from the genus Aplysina in southwest Puerto Rico. Caribbean Journal of Science 54: 57-76

*González-García María del Pilar, García-Sais JR, García-Moliner G, Schizas  NV (2024)  RAD-Seq derived SNPs reveal a weak population structure in the commercially important deep-sea queen snapper in Puerto Rico. Marine Life Science & Technology.


*Rodriguez Matos LR , Andras JP, E Weil, NV Schizas (2023) Genetic Connectivity among Gorgonia ventalina Linnaeus, 1758 (Cnidaria: Octocorallia: Gorgoniidae) Populations in the Caribbean Sea. Life: The Excitement of Biology. (10)3: 152-202.

*Cruz Ramos MA and NV Schizas (2023). Population structure of the fireworm Hermodice carunculata in the wider Caribbean, Atlantic and Mediterranean Sea. Journal of the Marine Biological Association, U.K. DOI: 10.1017/S0025315422001114

*Gonzalez Colmenares GM, Gonzalez Montes AJ, Harms-Tuohy CA, Schizas NV. 2023. Using eDNA sampling for species-specific fish detection in tropical oceanic samples: limitations and recommendations for future use. PeerJ 11:e14810 


Majeske A, AJ Mercado Capote, A Komissarov, A Bogdanova, NV Schizas, SO Castro Marquez, K Hilkert, W Wolfsberger, TK Oleksyk​ (2022) The first complete mitochondrial genome of Diadema antillarum (Diadematoida, Diadematidae). Gigabyte

*Báez Rodríguez N, Harms-Tuohy C, NV Schizas (2022). Using DNA barcoding to identify seafood fraud in Puerto Rico. Caribbean Journal of Science. 52(2) : 331-352

García-Urueña R, Kitchen SA, Schizas NV (2022) Fine scale population structure of Acropora palmata and Acropora cervicornis in the Colombian Caribbean. PeerJ10:e13854

*Laverde-Castillo JA, M Alfaro, E Weil, NV Schizas (2022) Polychaetes (Annelida, Polychaeta) associated with Mesophotic Coral Ecosystems in Puerto Rico and the U.S. Virgin Islands.  Caribbean Journal of Science 52: 82–107.

Schizas NV, H-U Dahms, P Kangtia, A Galindo Estronza. PHC Corgosinho (2022) Comment on Bang et al. Two New Species of the Genus Longipedia Claus, 1863 (Copepoda: Harpacticoida: Longipediidae) from Korea, with an Update and a Key to Species. Diversity, 2021, 13, 590

Diversity 14, 200. d14030200


Appeldoorn RS, DL Ballantine, M Carlo, JJC Motta, M Nemeth, HJ Ruiz, NV Schizas, CE Sherman, E Weil, PM Yoshioka (2021). Intra-annual Variation in Mesophotic Benthic Assemblages on the Insular Slope of Southwest Puerto Rico as a Function of Depth and Geomorphology.  Frontiers in Marine Science. doi: 10.3389/fmars.2021.732926

*García-Hernández JE, E Tuohy, D Toledo, C Sherman, NV Schizas, E Weil (2021). Detrimental conditions affecting Xestospongia muta across shallow and upper-mesophotic coral reefs off the southwest coast of Puerto Rico.  Diseases of Aquatic Organisms 147: 47–61 DOI:

*Ortiz-Gonzalez IC, *RE Rivera-Vicens, NV Schizas (2021)Description of four Millepora spp. transcriptomes and their potential to delimit the Caribbean fire coral species. Marine Genomics

*García-Hernández JE, NV Schizas (2021). The ‘anemone’ shrimp Periclimenes rathbunae perched on the stony coral Mussa angulosa in Puerto Rico. Marine Biodiversity 51:25 (pdf)

*Veglia AJ, Milford CR, NV Schizas (2021) Isolation and genotyping of novel T4 cyanophages associated with diverse coral reef invertebrates. Coral Reefs

*Hammerman NM, SM Williams, AJ Veglia, JE García-Hernández, NV Schizas, JC Lang (2021) Cladopsammia manuelensis sensu Cairns, 2000 (Order: Scleractinia): A new distribution record for Hispaniola and Puerto Rico. Cahiers de Biologie Marine 62: 1-10.

*González-García M del Pilar, NV Schizas, MV Concepción-Torres, and CE Diez (2021Lepidochelys olivacea in Puerto Rico: Occurrence and Confirmed Nesting.  Marine Turtle Newsletter No. 162: 13-17 (pdf)


Pesic V, T Chatterjee, M, NV Schizas (2020) A new marine mite of the genus Litarachna Walter, 1925 from Guadeloupe, Caribbean Sea (Acari, Hydrachnidia, Pontarachnidae). Ecologica Montenegrina 38: 205-209 (pdf)

*Rodriguez Ferrer G, JJ Cruz Motta, NV Schizas, RS Appeldoorn (2020) Modeling the distribution of the common bottlenose dolphin, Tursiops truncatus off the southwest coast of Puerto Rico. Journal of Marine Systems DOI:  10.1016/j.jmarsys.2020.103371

Nieves-Rivera ÁM, JP Zegarra Vila, CE Figuerola Hernández, JE García Hernández and NV Schizas (2020) New Faunal Records and Recent Explorations of the Underwater Section of Cueva del Agua, Punta Los Ingleses, Mona Island (Puerto Rico). Life: The Excitement of Biology 8(1) DOI: 10.9784/LEB8(1)NievesRivera.01 (pdf)

Chatterjee T, Dovgal I, NV Schizas (2020).  Report of epibiont ciliates (Ciliophora) on harpacticoid copepods from Caribbean​ mesophotic reefs​

Cah. Biol. Mar. 61: 131-136. (pdf)

Boisnoir A, Pavaux A-S, NV Schizas, S Marro, T Blasco, R Lemée, P-Y Pascal (2020). The use of stable isotopes to measure the ingestion rate of potentially toxic benthic dinoflagellates by harpacticoid copepods.  Journal of Experimental Marine Biology and Ecology 524: 151285 (pdf)



Hadfield KA, Schizas NV, Chatterjee T, Smit NJ (2019) Gnathia bermudensis (Crustacea, Isopoda, Gnathiidae), a new species from the mesophotic reefs of Bermuda, with a key to Gnathia from the Greater Caribbean biogeographic region. ZooKeys 891: 1–16. (pdf)

*Rivera-García L, R Rivera-Vicéns, A Veglia, NV Schizas (2019) De novo transcriptome assembly of the digitate morphotype of Briareum asbestinum (Octocorallia: Alcyonacea) from the southwest shelf of Puerto Rico. Marine Genomics

Pesic V, T Chatterjee, M, NV Schizas (2019).  A Checklist of Pontarachnidae (Acari: Hydrachnidia) and notes on distributional patterns of the species. Zootaxa 4619 (3): 527–544 (pdf)

Appeldoorn RS, Alfaro M, Ballantine DL, Bejarano I, Ruíz HJ, Schizas NV, Schmidt WE, Sherman CE, Weil E. 2019. Puerto Rico. In: Loya Y, Puglise KA, Bridge TCL. (eds) Mesophotic Coral Ecosystems of the World. Springer Amsterdam. (pdf)

Goulet TL, MQ Lucas, NV Schizas (2019). Symbiodiniaceae Genetic Diversity and Symbioses with Hosts from Shallow to Mesophotic Coral Ecosystems. In: Loya Y, Puglise KA, Bridge TCL. (eds) Mesophotic Coral Ecosystems of the World. Springer Amsterdam (pdf)

*Garcia-Hernandez JE, *Hammerman NM, Cruz-Motta JJ,  Schizas NV. (2019) Associated organisms inhabiting the calcareous sponge Clathrina lutea in La Parguera, Puerto Rico.  Caribbean Journal of Science 49:239-254 (pdf)


Sukhikh N, Souissi A, Souissi S, A-C Holl, NV Schizas, V Alekseev (2018).  Life in sympatry: coexistence of native Eurytemora affinis and invasive Eurytemora carolleeae in the Gulf of Finland (Baltic Sea). Oceanologia (pdf)

Schizas NV (2018) Hurricane Maria and its Impact on the Zoological and Botanical Collections of the University of Puerto Rico.  Biodiversity Information Science and Standards 2: e26014 doi: 10.3897/biss.2.26014 (pdf)

​*Veglia A, Hammerman NM, Rivera Vicens RE, NV Schizas (2018). De novo transcriptome assembly of the coral Agaricia lamarcki (Lamarck’s sheet coral) from mesophotic depths in southwest Puerto Rico. Marine Genomics (pdf)

*García-Hernández JE, Sanchez PJ, Hammerman NM, Schizas NV (2018). Fish, coral, and sponge assemblages associated with altiphotic and mesophotic reefs along the Guanica Biosphere Reserve continental shelf edge, southwest Puerto Rico.  Frontiers in Marine Science doi: 10.3389/fmars.2018.00303 (pdf)

*Veglia A, Hammerman NM, Rivera Rosaly CR, Lucas M, Galindo Estronza A, Corgosinho PH, and NV Schizas (2018). Characterizing population structure of coral-associated fauna from mesophotic and shallow habitats in the Caribbean. Journal of the Marine Biological Association of the United Kingdom. doi:10.1017/S0025315418000413 (pdf)

Dahms H-U, NV Schizas, RA James, L Wang, J-S Hwang (2018). Marine hydrothermal vents as templates for global change scenarios. Hydrobiologia (pdf)

Corgosinho P,  TC Kihara, NV Schizas, A Ostmann, P Martínez Arbizu, S Ivanenko (2018).   Traditional and confocal descriptions of a new genus and two new species of deep water Cerviniinae Sars, 1903 from the Southern Atlantic and the Norwegian Sea: with a discussion on the use of digital media in taxonomy (Copepoda, Harpacticoida, Aegisthidae).  ZooKeys  766: 1–38 (2018) doi: 10.3897/zookeys.766.23899 (pdf)


*Hammerman NM, *RE Rivera-Vicens, MP Galaska, E Weil, RS Appledoorn, M Alfaro, NV Schizas (2017).  Population connectivity of the plating coral, Agaricia lamarcki from southwest Puerto Rico. Coral Reefs DOI: 10.1007/s00338-017-1646-x (pdf)

*Beltran DR, NV Schizas, RS Appeldoorn, C Prada (2017). Effective Dispersal of a Caribbean Reef Fish is Smaller than Current Spacing Among Marine Protected Areas. Nature Scientific Reports | 7: 4689 | DOI:10.1038/s41598-017-04849-5 1

*Rodriguez G, RS Appeldoorn, NV Schizas (2017). Abundance of the common bottlenose dolphin, Tursiops truncatus (Montagu, 1821), off the south and west coasts of Puerto Rico  (2017). Life: The Excitement of Biology 4:(4) DOI: 10.9784/LEB4(4)RodriguezFerrer.01 (pdf)

Corgosinho PHC, Schizas NV , Previattelli D, Falavigna da Rocha CE, Edinaldo Nelson dos Santos-Silva (2017).  A new genus of Parastenocarididae (Copepoda, Harpacticoida) from the Tocantins River basin (Goiás, Brazil), and a phylogenetic analysis of the Parastenocaridinae.  Zoosystematics and Evolution  93: (1) 167–187 | DOI 10.3897/zse.93.11602 (pdf)


*Ortiz-Gonzalez IC, RE Rivera-Vicens, NV Schizas (2016). De novo transcriptome assembly of the Hydrocoral Millepora alcicornis (branching fire coral) from the Caribbean. Marine Genomics (pdf)

Petrescu, I, T Chatterjee, NV Schizas (2016). New species and new records of Cumacea (Crustacea: Peracarida: Cumacea) from mesophotic reefs of Puerto Rico and US Virgin Islands, Caribbean Sea. Zootaxa 4199 (1): 001–078 (pdf)

*Galindo Estronza A, NV Schizas, M Alfaro (2016). Morphological and genetic species diversity in ostracods (Crustacea: Oligostraca) from Caribbean reefs. Marine Biodiversity DOI: 10.1007/s12526-016-0525-y (pdf)


Appeldoorn R, D Ballantine, I Bejarano, H Ruiz, NV Schizas, W Schmidt, C Sherman, E Weil (2016). Mesophotic coral ecosystems examined: La Parguera, Puerto Rico, USA, in: Baker, E.K., Puglise, K.A. and Harris, P.T. (Eds.). (2016). Mesophotic Coral Ecosystems:A lifeboat for coral reefs? The United Nations Environment Programme and GRID-Arendal, Nairobi and Arendal.,www.grida.no5 pp 45-49.  (pdf) 

*Harms-Tuohy CA, NV Schizas, RS Appeldoorn (2016). The use of DNA metabarcoding for stomach content analysis in the invasive lionfish (Pterois volitans) in Puerto Rico. Marine Ecology Progress Series doi: 10.3354/meps11738  (pdf) and (Supplement File)

*Lucas MQ, M Stat, M Smith, E Weil, NV Schizas (2016). Symbiodinium (ITS2) diversity in the coral host Agaricia lamarcki (Cnidaria: Scleractinia) among shallow and mesophotic reefs (20 - 70 m). Marine Ecology doi: 10.1111/maec.12367 (pdf) 

*Ricaurte M, Schizas NV, P Ciborowski, NM Boukli (2016). Proteomic analysis of bleached and unbleached Acropora palmata, a threatened coral species of the Caribbean. Marine Pollution Bulletin 107:224-232 doi: 10.1016/j.marpolbul.2016.03.068 (pdf) 

Corgosinho PHC, NV Schizas, M Alfaro (2016) A new species of Atergopedia (Copepoda: Harpacticoida: Novocriniidae) from a Caribbean mesophotic reef. Marine Biodiversity DOI: 10.1007/s12526-016-0446-9 (pdf) 

Petrescu, I, T Chatterjee, NV Schizas (2016). A new species of the genus Campylaspis (Crustacea: Cumacea) from the mesophotic reefs of St. John, US Virgin Islands. Caribbean Journal of Science 49: (1)17-26 (pdf)


Appeldoorn R, David Ballantine, Ivonne Bejarano, Milton Carlo, Michael Nemeth, Ernesto Otero, Francisco Pagan, Hector Ruiz, Nikolaos V. Schizas, Clark Sherman, Ernesto Weil (2015) Coral Ecosystems and Impacts from Anthropogenic Stress: a case study at Ponce, Puerto Rico. Coral Reefs DOI: 10.1007/s00338-015-1360-5 (pdf)

Pesic V, T Chatterjee, NV Schizas (2015). First record of Litarachna caribica Pesic et al. 2008 (Acari, Pontarachnidae) from the Pacific coast of Panama. Marine Biodiversity Records. Vol. 8: e85  (pdf) 

Guerra-Garcia JM, T Chatterjee, NV Schizas (2015). New genus and new species of Caprellidae (Crustacea: Peracarida: Amphipoda) from mesophotic coral ecosystems of Puerto Rico and St. Croix, Caribbean Sea. Zootaxa 4018 (1):080-096 (pdf) 

Schizas NV, H-U Dahms, P Kangtia, PHC Corgosinho, A Galindo Estronza (2015). A new species of Longipedia Claus, 1863 (Copepoda: Harpacticoida: Longipediidae) from Caribbean mesophotic reefs with remarks on the phylogenetic affinities of Polyarthra. Marine Biology Research DOI: 10.1080/17451000.2015.10 (pdf) 

*Beltran D, NV Schizas, Prada C, R Appeldoorn (2015). Isolation and characterization of twelve microsatellite loci to study connectivity in the yellow jawfish Opistognathus  autifrons. Conservation Genetics Resources DOI: 10.1007/s12686-014-0419-x  (pdf) 

*Merten WB, NV Schizas, MT Craig, RS Appeldoorn, DL Hammond (2015). Genetic structure and dispersal capabilities of dolphinfish (Coryphaena hippurus): implications for stock-based fishery management in the western central Atlantic. Fishery Bulletin 113:419-429 doi: 10.7755/FB.113.4.5 (pdf)


Senna AR, R Sorrentino, T Chatterjee, NV Schizas (2014). A new species of Boca Lowry & Stoddart, 1997 (Amphipoda: Lysianassoidea: Aristiidae) from a mesophotic coral ecosystem off Puerto Rico, Caribbean Sea. Zootaxa 3884 (5): 429-436 (pdf)

Petrescu, I, T Chatterjee, NV Schizas (2014). New records of Bodotridae (Crustacea: Cumacea) from Puerto Rico with description of three new species. Zootaxa 3873 (5): 526-540(pdf)

Schizas NV, H-U Dahms, M Ricaurte Chica*, J-S Hwang (2014). Population Genetic Patterns of the Copepod Calanus sinicus in the NW Pacific. Hydrobiologia 741: 153-165 DOI 10.1007/s10750-014-2011-5 (pdf)

Pesic V, T Chatterjee, M Alfaro, NV Schizas (2014) A new species of Litarachna (Acari, Hydrachnidia, POntarachnidae) from a Caribbean mesophotic coral ecosystem with notes on a littoral Litarachna sp. Zookeys (pdf)

Chatterjee T, Fernandez-Leborans G, NV Schizas (2014). Report of Thecacineta calix (Ciliophora: Suctorea) as epibiont on harpacticoid copepods from mesophotic reefs of Puerto Rico and St. John, U.S. Caribbean. Caribbean Journal of Science (pdf)

*DV Ruiz-Ramos, E Weil, NV Schizas (2014). Morphological and genetic evaluation of the hydrocoral Millepora species complex in the Caribbean. Zoological Studies 53:4 (pdf)

Petrescu I, T Chatterjee, NV Schizas (2014). Three new Nannastacidae (Crustacea: Cumacea) species from a Caribbean mesophotic ecosystem. Zootaxa 3765 (4): 360-370 (pdf)

*Mege P, Schizas NV, Garcia J, Hrbek T (2014). Genetic seascape of the threatened Caribbean elkhorn coral, Acropora palmata, on the Puerto Rico Shelf. Marine Ecology doi: 10.1111/maec.12135 (pdf)

Petrescu I, T Chatterjee, NV Schizas (2014) New species of Cumella (Crustacea: Cumacea: Nannastacidae) from mesophotic habitats of Mona Island, Puerto Rico, Caribbean Sea. Cahiers de Biologie Marine 55(2): 183-189.(pdf)


Sherman C, Appeldoorn R, Ballantine D, Bejarano I, Carlo M, Kesling D, Nemeth M, Pagan F, Ruiz H, Schizas NV, Weil E (2013) Exploring the mesophotic zone: Diving operations and scientific highlights of three research cruises across Puerto Rico and US Virgin Islands. In: Lang, MA & Sayer MDJ (ed.), Proceedings of the 2013 AAUS/AEDP Curacao Joint International Scientific Diving Symposium, Dauphin Island, AL: American Academy of Underwater Sciences, pp 297-312 (pdf)

Petrescu I, T Chatterjee, NV Schizas (2013) Two new species of the genus Cumella (Crustacea: Cumacea: Nannastacidae) associated with mesophotic reefs of Puerto Rico and St. Croix, Caribbean Sea. Cahiers de Biologie Marine 54: 257-262 (pdf) 

Corgosinho PHC, Schizas NV (2013) Archeolourinia, a new genus and species of Louriniidae Monard, 1927 (Copepoda, Harpacticoida) from a Caribbean mesophotic zone. Journal of Marine Biological Association of the UK. doi:10.1017/S0025315412001336 (pdf)


NV Schizas (2012) Misconceptions regarding nuclear mitochondrial pseudogenes (Numts) may obscure detection of mitochondrial evolutionary novelties. Aquatic Biology 17: 91-96 (pdf) 

Pesic V, Chatterjee T, Schizas NV (2012) A new species of Pontarachna (Acari, Hydrachnidia, Pontarachnidea) from a mesophotic coral ecosystem off Vieques Island, Puerto Rico, Caribbean Sea. Zootaxa 3440: 63-67(pdf) or if you do not have access to Zootaxa (pdf)

Petrescu I, T Chatterjee, NV Schizas (2012) New genus and new species of Cumaceans (Crustacea: Peracarida: Cumacea) from the mesophotic coral ecosystem of SW Puerto Rico, Caribbean Sea. Zootaxa 3476: 55-61.  (pdf)


S Chullasorn, H-U Dahms, K-W Lee, J-S Ki, N Schizas, P Kangtia, HG Park, and J-S Lee (2011) Description of Tisbe alaskensis sp. nov. (Crustacea: Copepoda) Combining Structural and Molecular Traits. Zoological Studies 50(1): 103-117 (pdf)

*H Torres-Pratts, T Lado-Insua, AL Rhyne, L Rodriguez-Matos, NV Schizas (2011). Two distinct, geographically overlapping lineages of the corallimorpharian Ricordea florida (Cnidaria: Hexacorallia: Ricordeidae). Coral Reefs DOI: 10.1007/s00338-010-0709-z. (pdf) 

*DV Ruiz-Ramos, EA Hernandez-Delgado, NV Schizas (2011). Population status of the long-spined urchin Diadema antillarum in Puerto Rico 20 years after a mass mortality event. Bulletin of Marine Science. 87: 113-127 (pdf) or if you do not have access to the journal (pdf) 


GC Fiedler, AL Rhyne, R Segawa, T Aotsuka, NV Schizas (2010). The evolution of euhermaphroditism in caridean shrimps: a molecular perspective of sexual systems and systematics. BMC Evolutionary Biology 2010, 10:297 (pdf)

*J Garcia Reyes, NV Schizas (2010). No two reefs are created equal: fine-scale population structure in the threatened coral species Acropora palmata and A. cervicornis. Aquatic Biology. doi: 10.3354/ab00254. (pdf)

*PJ Milligan, EA Stahl, NV Schizas, JT Turner (2010). Phylogeography of the copepod Acartia hudsonica in estuaries of the northeastern United States. Hydrobiologia doi: 10.1007/s10750-010-0097-y (pdf) or if you do not have access to the journal (pdf) 


Andrew L. Rhyne, Dong Zhang, J Lin, NV Schizas (2009). Not any two will do: DNA divergence and interpopulation reproductive compatibility in a simultaneous hermaphroditic shrimp, Lysmata wurdemanni. Marine Ecology Progress Series 388:185-195 (pdf)

Hans-U. Dahms, S Chullasorn, NV Schizas, P Kangtia, W Anansatitporn, Wan-Xi Yang (2009). Review of naupliar development among Tisbidae (Copepoda: Harpacticoida) Tisbe thailandensis from Thailand. Zoological Studies 48(6):780-796 (pdf) 

*H Torres-Pratts, MT Scharer and NV Schizas (2009) Genetic diversity of Chelonibia caretta, commensal barnacles of the endangered hawksbill sea turtle Eretmochelys imbricata from the Caribbean (Puerto Rico). Journal of the Marine Biological Association of the UK 89:(4) 719-725 (pdf)(Copyright of Cambridge University Press).

S Chullasorn, H-U Dahms, NV Schizas, P Kangtia (2009). Phylogenetic inferences of Tisbe Lilljeborg, 1853 (Copepoda, Harpacticoida) with a new species Tisbe thailandensis from Bangsaen Beach, Thailand. Hydrobiologia 627: 1-17 DOI 10.1007/s10750-009-9711-2 (pdf)


V Pesic, T Chatterjee, NV Schizas (2008). Marine water mites (Acari: Hydrachnidia: Pontarachnidae) from the Caribbean Sea, with description of one new species. Cahiers de Biologie Marine 49(3):253-259. (pdf)

*C Prada, NV Schizas, PM Yoshioka (2008). Phenotypic plasticity or speciation? A case from a clonal marine organism. BMC Evolutionary Biology 8:47 (Featured article in BMC Evolutionary Biology, February 29 2008) (pdf)


*H Torres-Pratts, NV Schizas (2007). Meiofaunal colonization of decaying leaves of the red mangrove Rhizophora mangle, in southwestern Puerto Rico. Caribbean Journal of Science, 43 (1): 127-137. (pdf). High resolution pictures of Figure 2 (here) and Figure 3 (here).


NV Schizas, TC Shirley (2006).  Apolethon hippoperus (Copepoda, Harpacticoida)- a new species from southeastern Alaska. Journal of Crustacean Biology, 26 (3): 341-354. (pdf)


Hans-Uwe Dahms, NV Schizas, TC Shirley (2005). Naupliar evolutionary novelties of Stenhelia peniculata (Copepoda, Harpacticoida) from Alaska affirming taxa belonging to different categorical rank. Invertebrate Zoology, 2(1): 1-14. (pdf)


Staton JL, BE Taylor, NV Schizas, R Wetzer, TC Glenn, BC Coull (2003). Mitochondrial gene diversity of Skistodiaptomus mississippiensis in impoundments of the Upper Coastal Plain near Aiken, South Carolina, USA. Archive fur Hydrobiologie 158: 215-231 (pdf)


Staton JL, Schizas NV, Klosterhaus SL, Griffitt RJ, Chandler GT, Coull BC (2002). Effect of salinity variation and pesticide exposure on an estuarine harpacticoid copepod, Microarthridion littorale (Poppe), in the southeastern US. Journal of Experimental Marine Biology & Ecology 278: 101-110. (pdf)

Schizas NV, Coull BC, Chandler GT, Quattro JM (2002). Sympatry of distinct mitochondrial lineages in a copepod inhabiting estuarine creeks in the southeastern USA. Marine Biology 140: 585-594 (pdf)


Schizas NV, Chandler GT,Coull BC, Klosterhaus SL Quattro JM (2001). Differential survival of three mitochondrial lineages in a marine copepod exposed to a mixture of pesticides. Environmental Science and Technology. 35: 535-538 (pdf) 

Steinbachs J, Schizas, NV, JWO Ballard (2001). Efficiencies of genes and accuracy of tree-building methods in recovering a known Drosophila genealogy. Pacific Symposium on Biocomputing 6: 606-617 (pdf) 

Staton JL, Schizas NV, Chandler GT, Coull BC, Quattro JM. (2001). Ecotoxicology and Population Genetics: The Emergence of 'Phylogeographic and Evolutionary Ecotoxicology'. Ecotoxicology 10:217-222 (pdf) 


Kovatch CE, Schizas NV, Chandler GT, Coull BC (2000). Tolerance and genetic relatedness of three meiobenthic copepod populations exposed to sediment-associated contaminant mixtures: Role of environmental history. Environmental Chemistry and Toxicology 19: (4) 912-919 (pdf) 


Schizas NV, Street GT, Coull BC, Chandler GT, Quattro JM (1999). Molecular population structure of the benthic copepod Microarthridion littorale along the Southeastern and Gulf coasts of the United States. Marine Biology 135: 399-405 (pdf) 


Schizas NV, Street GT, Coull BC, Chandler GT, Quattro JM (1997). An efficient DNA extraction method for small metazoans. Molecular Marine Biology and Biotechnology 6: 381-383 (pdf) 

Chandler, GT, Coull BC, Schizas NV and Donelan, TL (1997). A culture-based assessment of the effects of chlorpyrifos on multiple meiobenthic copepods using microcosms of intact estuarine sediments. Environmental Chemistry and Toxicology 16: 2339-2346 (pdf)


Schizas, NV and TC Shirley (1996). Seasonal changes in structure of an Alaskan intertidal meiofauna assemblage. Marine Ecology Progress Series 133: 115-124. (pdf)


Schizas, NV and TC Shirley (1994). Onychocamptus krusensterni (Copepoda, Harpacticoida, Laophontidae)- a new species from Krusenstern Lagoon, Alaska. Crustaceana 66: 227-239 (pdf)

Schizas, N. V., and T. C. Shirley (1994). Autecology of an intertidal Alaskan harpacticoid copepod, Apolethon sp.--- Hydrobiologia 292/293: 361--367. (pdf)

Schizas, NV and TC Shirley (1994). Elapholaophonte decaceros n gen and n sp (Copepoda, Harpacticoida) from the Philippines. Journal of the American Microscopical Society 113: 127-141(pdf)

Schizas, NV and TC Shirley (1994). Two new species of Laophontodes (Copepoda, Harpacticoida, Ancorabolidae) from McMurdo Sound, Antarctica. Zoologica Scripta 23: 205-216  (pdf)

Schizas, NV and TC Shirley (1994).  Acrenhydrosoma maccalli, (Copepoda, Harpacticoida) a new species from Auke Bay, Alaska. Crustaceana 67: 329-340 (pdf)


Shirley, TC, M Carls, JW Fleeger and N Schizas (1993). Meiofaunal recolonization experiment with oiled sediments: major meiofauna taxa. In: Exxon Valdez Oil Spill Symposium Abstract Book. February 2-5, 1993, Anchorage AK, USA. Pp 350-353 (pdf)

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